桑村哲生『生命の意味 進化生態から見た教養の生物学』(第8版)裳華房、1008年3月20日。
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巌佐庸・松本忠夫・菊沢喜八郎・日本生態学会編 『生態学事典』 共立出版、2003年、ISBN 4-320-05602-7
沼田眞編著 『生態学辞典 増補改訂版』 築地書館、1983年、ISBN 4-8067-2350-9
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