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^ Erwin Stresemann, Ornithologische Monatsberichte XLIII. 138 en 1935
^ Revis, H.C. et Waller, D.A. (2004). “Bactericidal and Fungicidal Activity of ant chemicals on feather parasites: an evaluation of anting behavior as a Method of Self-medication in Songbirds” (英語). The Auk 121 (4): 1262?1268. 
^ Power, E. E et D. C. Hauser (1974). ⇒“Relationship of anting and sunbathing to molting in wild birds” (英語). The Auk 91: 537-563. ⇒http://elibrary.unm.edu/sora/Auk/v091n03/p0537-p0563.pdf
^ Zack, S. (1978). “Head grooming behaviour in the praying mantis”. Anim. Behav. 26: 1107?1119. 
^ Mather, J. A.; Alupay, J. S. (2016). “An ethogram for Benthic Octopods (Cephalopoda: Octopodidae)”. Journal of Comparative Psychology 130 (2): 109?127. doi:10.1037/com0000025. 
^ Gonzalez-Navarrete, D.A.; Vergara-Ovalle, F.; Garcia-Andaluz, P.; Ayala-Guerrero, F.; Rosas, C.; Vazquez-Leon, P.; Paz-Trejo, D.B.; Sanchez-Castillo, H. (2022). “Specific Ethogram of the Mexican four-eyed octopus: Octopus maya”. bioRxiv (preprint). doi:10.1101/2022.10.10.511610. 
^ Crook, R.J. (2021). Behavioral and neurophysiological evidence suggests affective pain experience in octopus. 24. pp. 102229. doi:10.1016/j.isci.2021.102229. 
^ Kuo, T.H.; Sneddon, L.U.; Spencer, J.W.; Chiao, C.C. (2022). “Impact of Lidocaine on Pain-Related Grooming in Cuttlefish”. Biology (Basel) 11: 1560. doi:10.3390/biology11111560. 
^ Leschnik, Michael; Joachim Weikel, Karin Mostl, Sandra Revilla-Fernandez, Eveline Wodak, Zoltan Bago, Elisabeth Vanek, Viviane Benetka, Michael Hess, Johann G. Thalhammer (2 2007). ⇒“Subclinical Infection with Avian Influenza A H5N1 Virus in Cats”. Emerging Infectious Diseases 13 (2): pp. 243-247. doi:10.3201/eid1302.060608. pmc: PMC2725870. ⇒http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2725870/ 2010年9月18日閲覧. "Therefore, the most likely route of transmission for these cats is contagious fecal contamination of the hair and oral uptake during grooming." 
^ Aureli, F., van Schaik, C., & van Hooff, J (1989). Functional aspects of reconciliation among captive long-tailed macaques (Macaca fascicularis). American Journal of Primatology, 19, 39-51.
^ Lawick-Goodall, J. van. (1968). The behavior of free living chimpanzees in the Gombe Stream Reserve. Animal Behavior Monographs, 1, 161-311.
^ de Waal, F. (1989). Peacemaking among primates. Cambridge, MA: Harvard University Press.
^ Smuts, B., Cheney, D., Seyfarth, R., Wrangham, R., & Struhsaker, T. (1987). Primate Societies. Chicago: University of Chicago Press.
^ Schino, G.; Scucchi, S.; Maestripieri, D.; Turillazzi, P.G. (1988), “Allogrooming as a tension-reduction mechanism: a behavioral approach”, American Journal of Primatology 16: 43?50, doi:10.1002/ajp.1350160106 
^ Smuts, Barbara (1985), Sex and Friendship in Baboons, New York: Aldine Publications, ISBN 978-0-202-02027-3, https://books.google.co.jp/books?id=Vx_l76muFWMC&printsec=frontcover&dq=smuts+%22Sex+and+Friendship+in+Baboons%22&q=&redir_esc=y&hl=ja 2010年4月28日閲覧。 
^ Gumert, Michael D. (December 2007), “Payment for sex in a macaque mating market”, Animal Behavior 74 (6): 1655?1667, doi:10.1016/j.anbehav.2007.03.009 
^ Moore, D.; Angel, J.E.; Cheeseman, I.M.; Robinson, G.E.; Fahrbach, S.E. (1995), “A highly specialized social grooming honey bee(Hymenoptera: Apidae)”, Journal of Insect Behavior 8 (6): 855?861, doi:10.1007/BF02009512 
^ Bshary, R.; Schaffer, D. (2002), “Choosy reef fish select cleaner fish that provide high-quality service”, Animal Behaviour 63 (3): 557?564, doi:10.1006/anbe.2001.1923 
^ Spruijt, B.M.; Van Hooff, J.A.; Gispen, W.H. (1992), “Ethology and neurobiology of grooming behavior”, Physiological Reviews 72 (3): 825?852, PMID 1320764 
^ Kimura, R. (1998), “Mutual grooming and preferred associate relationships in a band of free-ranging horses”, Applied Animal Behaviour Science 59 (4): 265?276, doi:10.1016/S0168-1591(97)00129-9 
^ Wilkinson, G.S. (1986), ⇒“Social grooming in the common vampire bat, Desmodus rotundus”, Animal Behaviour 34 (6): 1880?1889, doi:10.1016/S0003-3472(86)80274-3, ⇒http://www.life.umd.edu/faculty/wilkinson/Wilk_AB86.pdf 
^ Nelson, H. (2006), Human mutual grooming: an ethological perspective on its form and function 
^ Nelson, H. (2007), Encoding and decoding mutual grooming: Communication with a specialized form of touch 

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